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doi: 10.1016/j.ccr.2012.02.014 6 BalakrishnanKStellrechtCMGeniniDAyresMWierdaWGKeatingMJet al

H + is here a simplified notation of the actual chemical structure of the aqueous proton (whether H 13 O 6 + , H 5 O 2 + , or H 9 O 4 + , this is a debate beyond the topic of this article and covered by others [78, 103])

When GHK complexes with copper, it silences copper's redox activity, preventing the generation of damaging hydroxyl radicals while still allowing copper to be delivered into cells for use as an enzymatic cofactor

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